| Year | Position | Institution |
| 1958 | Massachusetts Institute Of Technology 4 | |
| 1964 | Postdoctoral Fellow | Systematics Ecology Program 4 |
| 1965 | Systematics Ecology Program 4 |
| Year | Role | Course |
| 1964 | Postdoctoral Fellow/Research Associate | Systematics-Ecology Program 1964 |
| 1965 | Postdoctoral Fellow/Research Associate | Systematics-Ecology Program 1965 |
| Year | Subject | Role |
| 1958 | Research Assistants | |
| 1964 | Independent | |
| 1965 | Independent |
| Year | Location |
| 1964 | marine biological laboratory |
| 1965 | marine biological laboratory |