| Year | Position | Institution |
| 1955 | Harvard University 4 | |
| 1959 | Fellow | Harvard University 8 |
| 1960 | Junior Fellow | Harvard University 8 |
| 1961 | Yale University 8 | |
| 1965 | Associate Professor Of Biology | Yale University 4 |
| 1966 | Associate Professor Of Biology | Yale University 4 |
| 1967 | Associate Professor Of Biology | Yale University 4 |
| 1968 | Associate Professor Of Biology | Yale University 4 |
| 1969 | Associate Professor Of Biology | Yale University 4 |
| 1970 | Associate Professor Of Biology | Yale University 8 |
| 1972 | Yale University | |
| 1972 | Professor And Chairman | Department Of Biology Yale University 4 |
| 1973 | Professor And Chairman | Department Of Biology 4 |
| 1973 | Yale University 4 | |
| 1974 | Professor Of Biology | Yale University 4 |
| 1975 | Professor And Chairman | Department Of Biology Yale University 4 |
| 1976 | Professor Of Biology | Yale University 4 |
| 1979 | Yale University 4 | |
| 1980 | Yale University 4 | |
| 2003 | Yale University 4 |
| Year | Role | Course |
| 1955 | Student | Physiology 1955 |
| 1959 | Instructor | Physiology 1959 |
| 1960 | Instructor | Physiology 1960 |
| 1961 | Instructor | Physiology 1961 |
| 1970 | Special Lecturer | Experimental Invertebrate Zoology 1970 |
| 1973 | Associate Staff | Experimental Invertebrate Zoology 1973 |
| 2003 | Lecturer | Analytical And Quantitative Light Microscopy 2003 |
| Year | Subject | Role |
| 1959 | Independent | |
| 1960 | Independent | |
| 1961 | Independent | |
| 1965 | Independent | |
| 1966 | Independent | |
| 1967 | Independent | |
| 1968 | Independent | |
| 1969 | Independent | |
| 1970 | Independent | |
| 1972 | Independent | |
| 1973 | Independent | |
| 1974 | Independent | |
| 1975 | Independent | |
| 1976 | Independent Investigators | |
| 1979 | Independent | |
| 1980 | Other Research Personnel |
| Year | Location |
| 1973 | yale university |
| Year | Start Date | End Date | Subject | Role |
| 1972 | - | - | Friday Evening Lecturer |